Triceratium barbadense Greville, 1861a, T. brachiatum Brightwell, 1856, T. inconspicuum Greville, 1861b and
T. kanayae Fenner, 1984a, are among the most common diatoms reported worldwide from lower to middle Eocene
biosiliceous sediments. Due to complicated nomenclatural histories, however, they are often confused. A morphometric
analysis performed herein indicates that T. brachiatum is conspecific with T. inconspicuum, and that both
were previously often misidentified as T. barbadense. Triceratium barbadense sensu stricto is a distinct species
similar to Triceratium castellatum West, 1860. Triceratium brachiatum and T. kanayae are transferred herein
to a new genus, Fenneria, for which a close phylogenetic relationship with Medlinia Sims, 1998 is proposed.
A review of the geographic and stratigraphic distribution of Fenneria shows that the best constrained records of
its occurrences are found at DSDP Site 338, and ODP Sites 1051 and 1260. The ages of the base (B) and top (T)
of each species’ stratigraphic range are calibrated here to the Geomagnetic Polarity Timescale either directly or
inferred via correlation with dinocyst biostratigraphy. Latitudinal diachroneity of ~7 million years is documented
for F. brachiata, which disappears earlier in tropical and mid-latitude sites than in the northern high latitudes. These
observations, coupled with a preliminary compilation of the Chron C20n taxonomic composition of pelagic diatom
assemblages for Sites 338, 1051 and 1260, indicate that diatoms diversified palaeobiogeographically considerably
earlier than the Eocene−Oligocene Transition, as commonly believed. This study also emphasizes the importance
of the detailed examination of specimens from both museum collections and deep-sea cores as a step toward enhancing
the utility of Palaeogene diatoms in palaeoceanographic and palaeoenvironmental reconstructions.
Early Palaeocene through early Eocene silicoflagellate assemblages were examined from five southern subtropical
through subpolar deep-sea sites: DSDP Holes 208 and 524, and ODP Holes 700B, 752A, and 1121B. For each
site, the taxonomic composition of the silicoflagellate assemblage is documented in detail; Pseudonaviculopsis
gen. nov., Dictyocha castellum sp. nov. and Stephanocha? fulbrightii sp. nov. are proposed, along with several
new combinations. More importantly, however, these observations enable a considerable refinement to the existing
Palaeocene–Eocene silicoflagellate biostratigraphic zonation that for the first time uses datums calibrated to
the Geomagnetic Polarity Timescale. The Corbisema aspera Interval Zone occurs immediately above the K/Pg
boundary and is here described from Seymour Island. The Corbisema hastata Partial Range Zone extends from
near the K/Pg boundary to late early Palaeocene and has been observed in Hole 208. The Pseudonaviculopsis disymmetrica
Acme Zone occurs in Holes 208 and 700B. The Dictyocha precarentis Partial Range Zone, observed
in Holes 208, 700B, 752A and 1121B, is subdivided into D. precarentis, Naviculopsis primativa, N. cruciata
and Pseudonaviculopsis constricta subzones. The Naviculopsis constricta Partial Range Zone occurs in Holes
524, 700B, 752A and 1121B. This study is also the first to consider syn- and/or diachroneity in Palaeogene
silicoflagellate biostratigraphy.